In this first programme he examines the origins of the once dominant savannah theory, and looks at the offence, outrage, and hostility caused by the rival aquatic ape theory.
Amongst those taking part are Philip Tobias, Elaine Morgan and Daniel Dennett.
The final programme investigates the recent evidence which has begun to push the argument away from the savannah theory and towards the beach-based, aquatic ape theory.
The programme concludes with provocative new evidence heard for the first time.
With contributions from Michael Crawford, Stephen Cunnane, Marc Verhaegen, Elaine Eorgan and Erica Schagatay.